Typical puberula occurs mostly in upland woods of central and southern Appalachian regions; it is common in e. Va. close to Ky. (F, Cr, HW+, FNA 26, K). There is a rather incomplete coll. of this species from BELL (TENN): L. Pounds CG85-31.5, 24 Apr 1985, Cumberland Gap National Historic Park, Ridge Trail near Butcher Gap. Also, J. Kiser (pers. comm.) has recently collected it (for EKY) from LETC in Pine Mountain Wildlife Management Area at Scuttlehole Gap. U. puberula differs from sessilifolia in its styles, which are separate to well below the middle (versus only in the upper 1/3 or 1/4); ovaries, sessile (versus distinctly stipitate); leaves, bright green on both sides (versus glaucous beneath); stems, puberulent in lines (versus glabrous); and rhizomes, very short with clustered roots (versus elongate, with scattered roots).