Mapping here is provisional, partly due to the need for more systematic study of colls. that takes account of differences between 'adult' and 'juvenile' shoots. The plants mapped here might result from more physiological maturity, or from more productive, damp or shady growing conditions (FNA 26, W). However, they appear distinct in some cases (F, Gl). Var. hederifolia has a south-central range similar to typical bona-nox. But F described it as often climbing high in "rich or damp woods and wet thickets," while typical bona-nox occurs on "dry to moist sands of dunes, clearings, fields and thickets" or on calcareous soils. In Ky. hederifolia is recorded mostly from broader bottomlands in southwestern regions, and from rich calcareous soils of the Inner Bluegrass. Var. hederifolia has broadly ovate-deltoid, subcordate leaves, at least on fertile branches, with little or no lobing (versus deltoid to panduriform with distinct lobes, often mottled with pale patches), and with spiny teeth fewer to absent along margins. These plants have often been misidentified as rotundifolia or as hispida; see notes under var. bona-nox for general characters of the species.